Grizzly 399’s ashes were spread in the Pilgrim Creek area inside Grand Teton National Park. That was her home turf and probable birth place. She died at the age of 28, having raised 18 cubs. Grizzly 1058’s remains were also “put back into the park landscape,” according to the National Park Service.

Grizzly 1058, a male from Grizzly 399’s famed 2020 litter of quadruplets, was confirmed dead on Tuesday, after apparently being hit by a vehicle in